From the Lady Slipper Archive: Kentucky’s ‘Tropical’ Fruit, the Pawpaw

The Lady Slipper newsletter of the Kentucky Native Plant Society has been published since the Society’s founding in 1986. This is one of a series of reprints from past issues. This article, about North America’s largest native fruit, the pawpaw (Asimina triloba), found in every county of KY, first appeared in the fall of 2005, Vol. 20, No. 3. If you would like to see other past issues, visit the Lady Slipper Archives, where all issues from Vol. 1, No. 1, February 1986 to Vol. 34, No. 1, Winter/Spring 2019 (after which we moved to this blog format) can be found.

The author, John Thieret, left a huge legacy to the native plant community when he passed in 2005. “Kentucky has lost its most renowned American plant taxonomist of the 20th century. John W. Thieret, Professor Emeritus of Biological Sciences at Northern Kentucky University, retired Director of the Northern Kentucky University Herbarium, Associate Editor of Sida, Contributions to Botany, and Editor of the Journal of the Kentucky Academy of Science (JKAS) passed away on 7 December 2005, at Alexandria, Kentucky.”

To learn more about this giant of Kentucky Botany, read the articles and tributes to him in the Winter 2005/Spring 2006, Vol. 20, No. 1, of the Lady Slipper archives.

Kentucky’s ‘Tropical’ Fruit,
the Papaw

by John Thieret, NKU

pawpaw fruits
Photo: Ellwood J.Carr, from the collection
of the Pine Mountain Settlement School

A visit to a fruit/ vegetable market in the tropics is a great experience. All sorts of plant products that we in the temperate zones do not recognize are there. Among these are fruits of the Annonaceae, the custard-apple family, including the bullock’s-heart, cherimoya, guanabana, sweetsop, and soursop. These are unknown to most people in our part of the world, but we do have a member of the Annonaceae that does NOT grow in the tropics, our papaw, Asimina triloba. This is a shrub or small tree, which, as I have seen it, never exceeds perhaps 20 feet in height and 6 inches in trunk diameter, although there are reports of individuals 50 feet tall and with a trunk 2 feet in diameter, truly a mega-papaw.

A common enough plant, the papaw thrives in rich woods over much of eastern U.S. from northern Florida to far eastern Texas, then north to New York, far southern Ontario, Michigan, Iowa, and southeastern Nebraska. It grows throughout Kentucky, almost certainly in every county.

Although some papaw enthusiasts wax ecstatic over the fruits, papaws are not everyone’s favorite. This divergence in appreciation stems from, first, natural differences in fruits from different trees and, second, differences in people’s taste buds. I have found fruits from some trees not worth the effort of trying to get them down from the branches. But other trees can produce fruits that I’d describe as almost excellent. The best papaws I ever tasted were in southern Illinois on a rather cool, almost frosty fall morning. Yes, quite worthwhile. The Indiana poet James Whitcomb Riley described, in hoosier dialect, the gustatory experience:

And sich pop-paws! Lumps a’ raw
Gold and green,—jes’ oozy th’ough
With ripe yaller—like you’ve saw
Custard-pie with no crust to.

Another assessment of the taste, by an Indiana lad, is included in Euell Gibbons’ book Stalking the Wild Asparagus: “They taste like mixed bananers and pears, and feel like sweet pertaters in your mouth.” I’ll second that, at least for a good papaw.

Long before Europeans began their assault on the North American continent,the indigenous peoples, along with various animals—possums, raccoons, squirrels, and skunks—sought the fruit. The first Europeans to see it—some 450 years ago—were De Soto and his entourage. They wrote of it, mentioning its “very good smell and excellent taste.” About 200 years later the plant was introduced into cultivation by Europeans who brought seeds to England. Then in1754 the first illustration of the papaw appeared in Catesby’s Natural History of Carolinas (see right). Lewis and Clark, in the early 19thcentury, found the fruits to be welcome additions to a meagre diet. To this day, the fruits are collected and used by country people and by city dwellers who like to eat their way through the landscape.

As for ways to use the fruits, first and foremost they can be eaten out of hand. As they ripen, they change from green to brown or nearly black, then looking not especially appetizing (recalling ripe plantains). The fruit pulp, creamy and sweet, contains several large,flattened, brown seeds. One of my friends made a necklace for his wife from the seeds. Better, I guess,than one made from finger bones.

Enthusiasts use the fruit for pies, puddings,marmalade, bread, beer, and brandy. I’ve tasted papaw bread and found it OK. Barely. I once tried to make papaw bread—I’ll say no more about that dismal experience. (The persimmon bread I attempted was no better.)

On a few occasions I have seen the plant grown as an ornamental. With its large, somewhat drooping leaves, it is rather attractive. The maroon flowers,which bloom in spring when the leaves are still young and covered with rusty down, are not all that conspicuous, and the fruits—well, my experience has been that papaw plants in cultivation as lawn specimens just do not make many fruits. As a matter of a fact, I have always noted that, even in the wild,the fruits are not abundantly produced. Maybe I just was not at the right place at the right time. The plants seem to require cross pollination, which is a disadvantage to those who would use them as ornamentals and, at the same time, would like some fruits.

If you have never tried one of the fruits, head for the woods in the autumn and attempt to find one. Maybe someone you know can help you. Even if you do not find the fruit much to your liking—maybe you will,maybe you won’t—you will have had a new gustatory experience.

For many years attempts have been made by horticulturists to ‘improve’ the papaw and make it into a commercially viable fruit. Their efforts notwithstanding, the fruit remains a Cinderella. On only one occasion have I seen papaws for sale: at a roadside farmer’s stand in southwestern Ohio among a fine display of squashes of a dozen kinds. Breeding and selection work has been carried out in several places, notably at Kentucky State University where about 1700 papaw trees grow in KSU’s 8-acre experimental farm and where the PawPaw Foundation is headquartered. Once, in Pennsylvania, I saw a papaw orchard of maybe 50 trees. I wish now that I had stopped and spoken with the orchard’s owner.Perhaps, with continued efforts at breeding and selection, papaws might some day be common items in our temperate fruit and vegetable markets, as common even as are the annonaceous cousins of Asimina triloba in markets of the tropics. This is the goal toward which papaw enthusiasts and breeders are striving.

From the Lady Slipper Archive: 2005 Wildflower of the Year, SHOWY GOLDENROD (Solidago speciosa)

The Lady Slipper newsletter of the Kentucky Native Plant Society has been published since the Society’s founding in 1986. This is one of a series of reprints from past issues. This article, about one of Kentucky’s loveliest goldenrods, first appeared in the spring of 2005, Vol. 20, No. 1. If you would like to see other past issues, visit the Lady Slipper Archives, where all issues from Vol. 1, No. 1, February 1986 to Vol. 34, No. 1, Winter/Spring 2019 (after which we moved to this blog format) can be found.

The author, Mary Carol Cooper, left a huge legacy to the native plant community when she passed in 2016. In almost every native plant gathering, her name is mentioned and a moment is given over to appreciate her knowledge, which she freely shared. Her passion led many of us to our love of natives; she was a mentor and friend to many of us.

2005 Wildflower of the Year
SHOWY GOLDENROD (Solidago speciosa)

By Mary Carol Cooper
Salato Native Plant Program Coordinator
Salato Wildlife Education Center

Showy goldenrod (Solidago speciosa)
Photo by Tom Barnes

Wildflower enthusiasts from all across the state have selected Showy Goldenrod (Solidago speciosa) as the Salato Native Plant Program’s Wildflower of the Year for 2005. The Wildflower of the Year is chosen based on the number of nominations it receives and how well it fits the established criteria; must be native to Kentucky, common and widespread across the state, seeds must be readily available, must be easy to grow, and must have good wildlife value.

Showy Goldenrod is a hardy perennial that grows 2 to 6 feet tall, depending on where it is planted. It is a rather showy species with stout, smooth, reddish stems and smooth, deep green leaves that are 4 to 10 inches and not toothed. It grows in rich thickets, woodland openings, fields, and prairies. It likes average to well drained soil and grows in sun to partial sun. It has dense upright pyramidal flower clusters. Each flower head has 6 to 8 rays. Showy Goldenrod blooms in late in the summer (August to September) and is wonderful as a late summer nectaring source for bees, butterflies and hummingbirds. It also provides food for several species of songbirds such as the Goldfinch, Junco, Pine Siskin, Song and Tree Sparrows.

Goldenrods are insect pollinated and their pollen is heavy and sticky. Therefore their pollen is never in the wind, so contrary to popular belief, this is not the plant that has always been blamed for causing hay fever. It is ragweed that causes all the misery! Ragweed blooms at the same time and is wind pollinated. I’ve enjoyed watching more and more floral designers use goldenrods in their arrangements and wonder how many people are aware that their lovely bouquet is full of the “dreaded goldenrod”.

Goldenrod is truly a North American flower. There are approximately 125 species in North America and more than 30 of these are native to Kentucky. Since the State Flower is Solidago ssp. this must mean that we have 30 State Flowers! Two of out native goldenrods, White-Haired Goldenrod and Short’s Goldenrod are on the Federally endangered species list.

Showy Goldenrod makes a nice background or midground plant in a sunny perennial garden. Establish this plant at the very rear of the garden or in the very middle of a circular or oval garden. Allow 3 feet between plants as this species grows into large clumps very fast. They can be divided every year or so and given to friends and neighbors. Nice companion plants are Ironweed, Great Blue Lobelia and New England Aster. Plants naturalize quickly on dry sunny banks. The cuttings are outstanding in arrangements.

The genus name Solidago comes from the Latin word that means “to make whole” or “to heal”, a name chosen because of medicinal power the plant was believed to have. The Native Americans used this plant for many things including ridding people from pain and evil spirits. One Goldenrod superstition says that he who carries the plant will find treasure, therefore, Goldenrod is the symbol for treasure and good fortune.

Goldenrod seeds and plants are available from many native plant nurseries. It is also very easy to propagate either by seeds, or division. Sow seeds thickly in outdoor seedbeds early in the fall or sow stored seed later in a flat indoors or in a cold frame. Transplant when there are 3 to 4 leaves. When the roots fill the pot, transplant in the garden after the last frost date. Collect seeds in late September or October. Cut off seed heads and put them upside down in a large paper bag. Let them dry for up to a week and then shake them in the bag and put the seeds in a sealed container.


Citizen science opportunity: have you seen falcata alfalfa?

By Jonathan O.C. Kubesch, University of Tennessee—Knoxville

Falcata alfalfa (Medicago sativa ssp falcata) is a stunning subspecies of the purple-flowered alfalfa commonly seen in the eastern United States (Figure 1; USDA-NRCS Plant Materials Program, 2015). Falcata alfalfa was introduced to South Dakota from Siberia by an eager professor in the early 19th century, however, the subspecies has appeared in the Kentucky flora in the time since (Shaw et al., 2020; Smith 1997). Falcata alfalfa improves the quantity and quality of forages for rangeland with tradeoffs for native species richness in South Dakota (Xu et al., 2004). The Kentucky collections may have come in seed bagged from elsewhere, especially as seed production shifted from the region to the Willamette Valley of Oregon (Figure 2).



Figure 1. Falcata alfalfa. Courtesy of Bing Creative Commons (accessed August 19, 2020).

Falcata alfalfa’s use in rangelands presents a question: why alfalfa has not naturalized into the South? Given it has historically been found in cultivated and disturbed environments, has the species formed any stable populations outside of agricultural use? And given falcata’s success in the High Plains, why hasn’t the species been used in the South?

Figure 2. The Willamette Valley of Oregon. July 10, 2020.
Figure 3. Falcata alfalfa (USDA PI 631577) growing on a misting bench in Knoxville, Tennessee. August 19, 2020.

Falcata is promising given its performance in low fertility conditions (USDA-NRCS Plant Materials Program, 2015). A research project at the University of Tennessee is investigating the horticultural performance of falcata alfalfa in the Southeast. Seed from the United States Department of Agriculture—by way of Canada and originally from eastern Italy—is being grown to study the plant in the eastern United States (Figure 3; USDA PI 631577). Concerns about introducing new plant species into the Southeastern flora prompted this present article. A fair number of plants introduced to the eastern United States have gradually moved outside of agricultural fields, such as tall fescue (Schedonorus arundinaceus), but the subspecies of alfalfa have not escaped and persisted for extended periods in the wild. 

The Tennessee-Kentucky Plant Atlas is a helpful tool for this sort of research (Shaw et al., 2020). Documenting the localities and conditions that falcata alfalfa has been found in the native flora should expound on these questions and theories regarding the species and subspecies persistence in the Southeast. Through the Atlas we can tell that escaped alfalfa does not necessarily take over native habitat in Kentucky and Tennessee in the same way that alfalfa has persisted in the High Plains of South Dakota. However, the Atlas is limited to collections made before 2002 (Shaw et al., 2020). With that in mind, citizen scientists are critical to documenting falcata alfalfa in Kentucky in 2020. The species should be seen where the purple flowered alfalfa occurs, such as disturbed areas and pastures.

Continue reading Citizen science opportunity: have you seen falcata alfalfa?

iNaturalist Tutorial

By Nick Koenig

Ever run across a flower in the forest and want to know what it is immediately? The app “iNaturalist” is a great tool that can serve as an excellent addition to field guides to help a plant lover come to a correct identification!

Video on How to Use iNaturalist

Associate Editor – Nick Koenig
Nick Koenig is a senior Biology Major with a Botany Concentration at Eastern Kentucky University.  At EKU, he serves as the Co-President of the student environmental group, Green Crew, works as a Biology Tutor, and volunteers in the Greenhouse. He fell in love with plants through gardening and the Kentucky State Fair but has continued with his passion through research at EKU.

Pollinators of Native Plants: A Book Review

Written by Heather Holm; Published by Pollination Press, LLC

Reviewed by Susan Harkins

I’m fond of saying that my favorite flower is the one I’m looking at. Similarly, my favorite book is the one I’m currently reading. However, when I say that Heather Holm’s Pollinators of Native Plants is one of my favorite books, I mean it. I keep just a few reference books close at hand, and this book is one of them.

It’s the most comprehensive, photo-filled book I’ve found on identifying the native pollinators that depend on our native flora. Every plant has a two-page spread that contains what you need to know about that plant to determine if it’s a good fit for your property and a full page or more of all the native pollinators it will attract. You get a full picture.

Its use goes beyond identifying pollinators: it’s one of the first books I pull out when wanting information about a particular plant, with or without consideration of its pollinators. However, when I want to identify a pollinator, it’s my go-to book. Often, I’m looking for host plants. This book supplies that information plus clear pictures of the pollinator in larvae form, which is invaluable when determining if I have an invading force in the yard or I’m seeing normal wear and tear by native beneficials.

The largest part of the book comprises the two-page spreads of 65 native plants. A few chapters in the beginning offer general advice from anatomy to ecothreats. At the end of the book, you’ll find visual botany glossaries, charts, garden designs, and more.

Pollinators of Native Plants is an outstanding reference on native trees and flowers that support our native pollinators. The detailed yet friendly layout make it easy to apply the information to your own property.

Heather Holm currently lives in Minnesota where she assists with native been research projects, the most recent being a study on native pollinators and cultivated blueberries. As a community supporter, she helps restore city-owned property for pollinators and people. You can keep up with Heather’s speaking engagements here: https://www.pollinatorsnativeplants.com/.


Susan Harkins is the Managing Editor of The Lady Slipper.

Presidents message

Cumberland Pitch Pine Barrens of Pine Mountain, a rare community in Kentucky

Hello fellow native plant enthusiasts.  It’s been roughly 6 months since the pandemic took hold of our daily lives.  Plans were put on hold.  Refocused efforts on health and safety of our families and friends immediately became priority.  KNPS had started out the year with a lot of momentum with our strategic planning meeting, and I would like to personally thank our board member Jeff Nelson for leading this effort.  Our committee members were planning hikes, symposiums, seed exchanges, forming an official plant conservation alliance, and bringing back our native plant stewardship program.  But these plans quickly had to be canceled or postponed out of caution for our members and friends.  So much uncertainty affected everyone and it does not seem to be getting any better at this time.  Life is precious and we only have one chance to live it.  It’s okay to switch gears in order to remain focused on what is fundamentally most important.  Despite all the unexpected events of this year, our online newsletter continued with some really great articles.  I want to personally thank Susan Harkins, our editor in chief, for all her work organizing and publishing The Lady Slipper along with coeditor and EKU student Nick Koenig. 

We are now beginning to regroup and plan for a virtual meeting this fall.  In the meantime, I know many of you all have been getting outside and continuing to find solace and comfort in the constant yet ever changing natural world, where spring fades to summer, fall, and the inevitable winter,  always changing, yet always there for us to witness all of the amazing diversity we have outside our back doors.  Seek out diversity of the plants in our natural areas, as well as the company you keep.  It’s important.  We will all come together again eventually, but until then let’s navigate this virtual world. 

Join our Facebook and Instagram groups.  Join the discussions.  Don’t forget to document your observations on our naturalist projects for documenting native plants on roadsides and our Botanists Big Year 2020, https://www.inaturalist.org/projects/kentucky-roadside-native-plants and https://www.inaturalist.org/projects/kentucky-botanists-big-year-2020.  If you are interested in helping with any of our ongoing plant conservation projects, please contact us to find ways to plug you into some of our ongoing projects that we coordinate through the Office of Kentucky Nature preserves.   

Tara R. Littlefield, KNPS president and Kentucky Plant Conservation Coordinator

Above are a few of the interesting plants and communities I have worked on this year, from remnant grasslands in our natural areas, powerline and roadsides, to seeps/bogs and orchid restoration, and forest biodiversity surveys. Kentucky is such a beautiful state with so much diversity! T.R. Littlefield

From The Lady Slipper Archives: New England Aster: 2010 Wildflower of the Year

The Lady Slipper newsletter of the Kentucky Native Plant Society has been published since the Society’s founding in 1986. With this article, we will begin to occasionally feature an article from a past issue. This one, about one of Kentucky’s loveliest natives, the New England Aster, first appeared in the summer of 2010, Vol. 25, No. 2. If you would like to see other past issues, visit the Lady Slipper Archives, where all issues from Vol. 1, No. 1, February 1986 to Vol. 34, No. 1, Winter/Spring 2019 (after which we moved to this blog format) can be found.

The author, Mary Carol Cooper, left a huge legacy to the native plant community when she passed in 2016. In almost every native plant gathering, her name is mentioned and a moment is given over to appreciate her knowledge, which she freely shared. Her passion led many of us to our love of natives; she was a mentor and friend to many of us.

New England Aster: 2010 Wildflower of the Year

By Mary Carol Cooper, Salato Center Native Plant Program Coordinator

The Wildflower of the Year is chosen based on the number of nominations it receives and this year more wildflower enthusiasts statewide voted than ever before! They chose New England Aster (Symphyotrichum novae-angliae) as the Salato Native Plant Program Wildflower of the Year for 2010. Aster comes from the Greek word for “star.” It describes the star-like form of the flower. Other familiar words using “aster” are astronomy, astrology and astronaut. According to Greek legend, the aster was created out of stardust when Virgo (the maiden Astraea, goddess of innocence and purity) looking down from heaven, wept. Asters were scared to all the gods and goddesses and beautiful wreaths made from the blossoms were placed on temple altars on very important festive occasions. Known in France as “eye of Christ” and in Germany as starworts, asters were often burned to keep away evil spirits. A hodgepodge of asters was thought to cure the bite of a mad dog. Shakers used the plant to clear their complexions and ancient Greeks used it as an antidote for snakebites and to drive away snakes. Virgil believed that boiling aster leaves in wine and placing them close to a hive of bees would improve the honey. Native Americans found many uses for asters, from treating skin rashes and earaches to stomach pains and intestinal fevers. Nerve medicines and cures for insanity were made from some asters and others were eaten as food. Some were smoked in pipes as a charm to attract game, especially deer. Today there are no medical uses for asters.

New England Aster, photo by Thomas Barnes

The genus Aster has recently undergone a name change due to close study using DNA testing and other techniques. There are about 150 flowering plants in North America traditionally placed in the aster genus. About 50 of them are considered common and widespread. Now there is only one species left with the name Aster. The other species have been given several tongue-twisting generic names. For the botanist, renaming of the asters brought accuracy and order. For the layperson, it removed some wonderfully colorful names and replaced them with unspellable and unpronounceable names! Aster novae-angliae was translated as “star of New England” and now as Symphyotrichum novae-angliae, it is literally “fused hairs of New England.” The word Symphyotrichum was created in 1832 to describe the hairs on the seeds of a European plant.

New England Aster is an erect perennial that grows to a height of 2’ to 6’ tall with a stout root crown or thick, short rhizome and clustered stems, usually with spreading hairs. The leaves are alternate, sessile, entire, lanceolate, 1” to 4” long with pointed lobes at the base that conspicuously clasp the stem. The ray flowers range from violet, rose, or magenta and are very showy. The disk flowers are yellow. This aster is one of our largest and showiest asters. There can be from 40 to 80 ray flowers on a head! These asters bloom from August to October and are a critical late-season nectar plant for butterflies, especially the Monarch, that stock up for their long migration to Mexico. New England Asters are found in mesic to wet open woods and fields across Kentucky. They prefer average to moist soil and full sun. Not only are the New England Asters critical for Monarch butterflies, it is the host plant for the Pearl Crescent and one of the host plants for the Saddleback Caterpillar Moth. Several game birds, including the wild turkey, a few songbirds, including the tree sparrow, and small mammals, such as the chipmunk and white-footed mouse, feed on the leaves and seeds. Work plantings of New England Aster into your fall landscape. Use them singly or in small groups in the rear of a sunny border. They look beautiful with our native sunflowers, goldenrods, mistflower and rose mallow. They are also perfect for rain gardens as they thrive on moist to dry soil. They are easy to naturalize in roadside ditches, road banks, and open grassy areas. A sunny site where soil remains moist throughout the season is also ideal. Asters have always been recognized as decorations. The flowers of most species last several days after being picked and put into vases, so what better than New England Asters in beautiful fall arrangements along with other fall bloomers.