Associate Editor Jonathan Kubesch earns his PhD from Virginia Tech

This photo shows editor Jonathan Jonathan Kubesch and family.
Dr. Jonathan Jonathan Kubesch and family.

Jonathan Omar Cole Kubesch, an associate editor and contributor to The Ladyslipper, graduated with his PhD in Crop, Soil, and Environmental Sciences from Virginia Tech on December 15, 2023.

Jonathan has been an advocate for native plants and ecosystems in unconventional spaces throughout the Midwest and Upper South. His doctoral work was part of a National Institute of Food and Agriculture grant regarding bee-friendly beef. Jonathan’s work focused on the establishment of complex native grass and wildflower mixtures in grazing systems to enhance livestock productivity and welfare as well as provide resources to native pollinators.

Jonathan’s dissertation abstract is presented below:

“Tall fescue grasslands are the dominant form of grasslands in the North American transition zone, however, warm-season forages provide necessary forage for livestock in the summer. Pollinators require steady forage availability in the same fashion as cattle. Native warm-season grasses (NWSG) and wildflowers (WF) could be incorporated into tall fescue (TF) grasslands to improve pollinator resources within agricultural landscapes without sacrificing livestock production. This balance of ecosystem services can be considered bee-friendly beef. Previous establishment experiments suggest that bee-friendly beef is a possibility, although establishment and persistence of NWSG-WF stands warrants additional research. Three small plot experiments were conducted to evaluate different planting methods designed to optimize establishment of NWSG-WF stands. Additionally, a grazing study examined whether NWSG-WF pasture enhancement could improve animal performance in TF grazing systems. The small plot experiments experienced extreme weed competition in the establishment year, although plots with higher NWSG content had fewer weeds over time. Floral production was comparable across treatment, which suggests lower WF seeding rates can generate comparable pollinator resources. Major findings from these experiments suggested that spatial or temporal separation of NWSGWF was unnecessary to generate successful establishment, varying the ratio of NWSG-WF in seed mixtures produced similar establishment outcomes, and that adding companion crops to NWSG-WF mixtures did not improve establishment success appreciably. In the grazing experiment, animal performance was improved in the biodiverse grazing systems when heifers 3 had access to the NWSG. Biodiverse and shaded grazing systems modified animal behavior and reduced body temperature relative to control grazing systems, although only biodiverse grazing systems improved animal performance. Native WF species failed to establish within the biodiverse grazing system, but white clover provided all subsequent floral production. Establishing WF in pastures is challenging and warrants additional study on more effective weed control strategies and systems to ensure establishment success. Selecting WF for agronomic performance or breeding such plant material will be necessary to improve emergence and establishment dynamics. Ideally trait data can identify the optimal WF species for specific grazing systems. Biodiverse NWSG-WF can be highly effective at improving summer weight gain in beef cattle. Future studies should focus on finding a more optimal balance of NWSG, WF, and TF that can benefit beef cattle production, pollinator services, and ecosystem services regarding soil and water quality, as well as pasture resiliency to dynamic conditions.”

The Buzz on Kentucky’s Native Bees 

By Katie Cody, Office of Kentucky Nature Preserves

Our native plants need native pollinators; they support nearly 90% of the world’s flowering plant reproduction. This pollination is mainly carried out by insects.  

Pollinators are considered a keystone species because they are the glue that holds an ecosystem together; without these species, the ecosystem could collapse. In this way, pollinators are helping maintain the structure and function of our natural communities. Given their importance, the alarm has recently been sounded on their decline, which is happening worldwide. These declines are being driven by many factors, such as climate change, introduced species, agricultural intensification, land use change, and pesticide use, among others. 

Graphic credit: Virginia R. Wagner, 2021 in: Wagner DL, Grames EM, Forister ML, Stopak D. 2021. Insect decline in the Anthropocene: Death by a thousand cuts. Proc Natl Acad Sci U S A. 118(2): e2023989118 

Of the pollinating insects, bees are the star of the show. But when many people think about bees, their minds may immediately go to the European honey bee (Apis mellifera). However, when we talk about our native bees, it isn’t fair to include this species — there are no honey bees native to the United States. Honey bees are a predominantly managed species and have vastly different life histories than most of our native bees.  

It’s important to also acknowledge that honey bees can negatively impact our native bees by outcompeting them for forage foods, decreasing their forage rates. They can even exacerbate the spread of invasive plants, which can distract our native bees from our native plants. Our native flowering plants and diverse natural areas depend on native bee visitation and diverse native pollinators to persist, not honey bees.  

Continue reading The Buzz on Kentucky’s Native Bees 

Botanical Timeline for Kentucky, 2015 to 2022 

By Ron Jones, Foundation Professor Emeritus, Eastern Kentucky University with assistance from Tara Littlefield 

This botanical timeline is a continuation of the series that appeared in The Lady-Slipper in five installments from 2005 to 2008:  

The goal is to highlight major events in the history of Kentucky botany, including new discoveries, important publications, changes in agricultural developments, major changes or upgrades of herbaria, developments or changes for the Kentucky Native Plant Society, and the retirements or deaths of people that made major contributions to Kentucky botany. Much more has happened and will continue to happen; follow-up lists and updates are encouraged. 

2015—Flora of the Southern and Mid-Atlantic States Working Draft of 21 May 2015  

Alan S. Weakley University of North Carolina Herbarium (NCU) North Carolina Botanical Garden University of North Carolina at Chapel Hill. Area covered is indicated in map below. See update below for 2022. 

2015—Guide to the Vascular Plants of Tennessee, by the Tennessee Flora Committee (editors: E.W. Chester, B.E. Wofford, J. Shaw, D. Estes, and D.H. Webb) 

University of Tennessee Press. 813 pages. (from UT Press): The product of twenty-five years of planning, research, and writing, Guide to the Vascular Plants of Tennessee is the most comprehensive, detailed, and up-to-date resource of its kind for the flora of the Volunteer State, home to nearly 2,900 documented taxa. A team of editors, authors, and contributors not only provide keys for identifying the major groups, families, genera, species, and lesser taxa known to be native or naturalized within the state—with supporting information about distribution, frequency of occurrence, conservation status, and more—but they also offer a plethora of descriptive information about the state’s physical environment and vegetation, along with a summary of its rich botanical history, dating back to the earliest Native American inhabitants. Other features of the book include a comprehensive glossary of botanical terms and an array of line drawings that illustrate the identifying characteristics of vascular plants, from leaf shape and surface features to floral morphology and fruit types. Finally, the book’s extensive keys are indexed by families, scientific names, and common names.  

The result is a user-friendly work that researchers, students, environmentalists, foresters, conservationists, and anyone interested in Tennessee botany and the surrounding states will value for years to come. 

2015—Herbarium at Centre College in Danville transferred to EKU  

A collection of a few hundred vascular plant specimens remaining at the Centre College Herbarium was transferred to EKY. Heidi Braunreiter facilitated this transfer, as she was working on her MS thesis on the flora of Boyle County at the time. The collection had formerly been curated by Susan Studlar and focused on nonvascular plants. When Susan Studlar accepted a position at West Virginia University, she took her collection of over 2,000 (mostly) Kentucky mosses and hepatics with her, leaving only a small collection of vascular plants at the college. 

2015—White-haired goldenrod (Solidago albopilosa), one of Kentucky’s three endemic plants, is delisted by USFWS in September 2015. The discovery of new populations and stable long-term trends documented by KSNPC botanists Deb White, Nick Drozda, and Tara Littlefield led to the realization that many of the populations were stable and that the goals of the White-Haired Goldenrod Recovery Plan were met. 

2015—George P. Johnson passes away in December 

This is a photo of George Johnson

George Johnson received his undergraduate and master’s degrees at Western Kentucky University and published a flora on the vascular plants of Barren County. He received his Ph.D. from North Carolina State University, working on the chinkapin chestnuts. He taught at Lindsey Wilson College for four years, before accepting a position at Arkansas Tech University in 1990. In 2015, he was promoted to full professor and honored for 25 years of service. Throughout his career, George remained active in botanical collecting, herbarium work, mentoring students, and publishing articles, and was co-editor of the 2014 volume on the Atlas of the Vascular Plants of Arkansas. He was particularly interested in working to preserve and enhance the herbaria in Arkansas and in developing a digital database of the collections and was a key figure in working with other southeastern U.S. curators to build a database of all SEUS collections, which culminated in the SERNEC website. George was held in very high regard by his friends and associates; for more on his life and career, see the 2016 obituary (from which this entry was copied) written by his student and colleague Travis Marsico: J. Bot. Res. Inst. Texas 10(1):295–298.  

Continue reading Botanical Timeline for Kentucky, 2015 to 2022 

How to turn a milk jug into a mini greenhouse

By Emilie Grace Yochim

When winter creeps in with her cold, frosty air, and gloomy, shorter days, most gardeners can only think about one thing: next year’s flower garden. Well, at least, that’s what runs through my mind. How can I keep myself busy with my garden over the dreary days of winter? Two words: winter sowing.

Winter sowing is a fun, cost effective way to prepare for next summer’s native garden. Defined, winter sowing is a way to start seeds outdoors during the winter months. It is particularly useful for any seeds that require a period of cold, moist stratification to break dormancy and germinate. Another advantage of winter sowing is that you end up with plants that are hardier, and acclimated to the temperature swings. Also, you don’t need to start seeds indoors using fancy grow lights or an expensive greenhouse! 

When to start

When to start? Most winter sowers begin their project on the winter solstice. Typically, you will start with your seeds that need the longest period of time to break dormancy. Those would be the seeds that need 60 to ­120 days of cold, moist stratification. You can winter sow as late as March using seeds that need less time or seeds that need no special treatment. 

What you need

What do I need to winter sow? Well, seeds would be at the top of the list! What “new to me” native plants and grasses would I like to add? What areas need more fillers to support the other plants I already have? Which plants did the pollinators love that I want to increase in number to support them? These are all questions you can ask yourself as you decide on seeds to grow.

Where can I get native seeds to winter sow? Some reliable sources to purchase native seeds include Roundstone Native Seed Company and Prairie Moon Nursery. You can also find native seed swaps in several groups on Facebook in Kentucky, including mine Pollinators, Poof Heads & Native Plants in Kentucky, where you can get seeds for free or a minimal fee to cover shipping costs. 

I have my seeds, now what? The following materials are what you’ll need on hand:

  • Gallon milk jugs: The clear ones work best; recycle the caps as they aren’t needed; you can also use such things as two-liter soda bottles and rotisserie chicken containers.
  • Potting soil: I tend to use Miracle Gro Potting Mix. Use whatever brand you prefer, but don’t use any that are labeled “moisture control.”
  • Duct tape: For sealing jugs.
  • Sharpie marker: You’ll want to identify the seeds in each jug. You could also place a plant marker on the inside of your jug.
  • Box cutter: For cutting your jug.
  • Drill, hot glue gun, or anything sharp you can safely use to make drainage holes in your jug.

The process

With all your materials gathered, you’re ready to start:

  1. Using a drill, hot glue gun, or something sharp, make 6 to 8 drainage holes in the bottom of a milk jug. I go around the lower sides of the jug.
  2. Use your box cutter to slice your jug from side to side, leaving the handle attached, about halfway down. Don’t cut the entire top portion of the jug; leave a few inches intact. The top will act as a lid that you can open and close.
  3. Add 4 inches, or so, of wet potting soil into the bottom of the jug. 
  4. Sow seeds according to directions. For example, you’ll sow some seeds on the surface and plant others at specific depths.
  5. Close the lid, and tape the jug back together using duct tape.
  6. Label the jug with your Sharpie. 
This photo shows the different stages of a milk jug being transformed into a mini greenhouse for propagating seeds.

Once your jugs are filled and taped shut, place your jugs in an area with sun. Moisture (rain and snow) will enter through the top creating condensation (aka your greenhouse). You can spray the inside if things start to dry out. 

This photo shows numerous milk jugs heeled in for winter.

Now, sit back and let nature take its course! Once spring is here, and temperatures warm up, open your jugs. Be sure to water as needed. If we get a frost or freeze, put the lids back down or cover the jugs with a blanket. 

This photo shows a couple of milk jugs with small germinating plants.

Once seedlings are ready to transplant in late spring, take them out of their containers, separate them in “hunks,” and plant them directly in the ground. If they’re small and still need more growing time, use the same method, but place them in pots to grow until they’re large enough to put in the ground. If your seedlings are large enough to separate, I’ll use Asclepias incarnata (Rose milkweed) as example, then by all means separate them instead of using the “hunk” method. 

Winter sowing helps us look forward to spring with the anticipation of new life growing in our containers and the pollinators that will enjoy them in the summer. It is a highly addictive project; you have been warned! I winter sowed over 130 containers last year, and I can say I likely won’t do as many this year, but then my plant pals who know me will just laugh and say, “yeah right.”

And, as always, please remember to support your local native plant growers. We need them to be successful in making native plants more accessible to the public and retailers!

Editors’ Note: You can follow Emilie’s backyard restoration by reading Replacing Invasives With Natives Is A Work Of Love.


Emilie Grace Yochim

Emilie Grace Yochim is a homeschooling mom to two teens and has been married to Philip for 20 years. Since 2019, Emilie has developed a slight obsession with pollinators, poof heads (bantam Satin and Silkie chickens), and native plants.

She enjoys educating others about planting natives and identifying pollinators by sharing pictures and information about the native plants, chickens, and wildlife in her yard. She also enjoys making terrible memes and puns that she finds hilarious.

The Kentucky native plant swap, four years and going strong

By Anne Milligan

The Kentucky native plants/seed swap movement is now in its fourth year, and this is an update on our progress. As many of you know, our swaps began in 2019 in Louisville, when some of us realized that we needed a venue to share all the extra seeds and plants from our native habitats. As of August 2023, we have 10 regional swaps fanning out from Louisville in all directions and a Louisville citywide Facebook group of over 2,500 members.  

Our newest regional swap group is also a transitional moment for our greater swap network. A young lady in Somerset stepped up to host a swap group for the entire Pulaski County/Lake Cumberland area. She envisions that group as a hub for an expanding swap network independent of the Louisville one.

Our Louisville citywide “big swap” will take place at Jefferson Memorial Forest in Fairdale. The staff offered us the Horine Conference Center for free, and for that we are very grateful. Our big swap brings together the extra seeds accumulated from all the regional fall swaps that took place last November. Regional swap hosts will be present to help guide new folks on native gardening and to check for nonnative species, which we don’t want to spread around.

We’re happy to announce that this past year, many of us expanded into growing the earliest-blooming native species, or spring ephemerals, and we are excited to share those extra seeds in the years ahead at regional spring swaps.

Our goal is to increase the diversity of Kentucky native plant species, primarily by educating property owners/gardeners/landscapers via regional swap groups how to grow and share native species. We understand that, by doing so, we are joining  passionate groups of people all over the world who are helping to restore natural ecosystems essential for life on this planet. To keep up with the various swap events, please join our Louisville Citywide Facebook group by clicking here: https://www.facebook.com/groups/652100455295916

We encourage everyone to establish native plant swap events in every Kentucky county, administered by individuals who are passionate about native species. There has never been a more important time than now to use our own properties to restore native ecosystems, not just for ourselves but for all those who follow.

Editors’ Note: To learn how this group got started, read Follow A Growing Trend And Organize Your Own Seed Swap. Read a review of Anne’s book, “Let the Earth Breathe,” which she coauthored with her husband, Stephen Brown.


Anne Milligan is an artist, singer/musician, and landscape designer. She lives in Louisville, KY with her husband, author and historian Stephen A. Brown.

Kentucky Pollinator Protection and Monarch Conservation group hold stakeholders meeting 

By Michaela Rogers, Kentucky Department of Fish and Wildlife Resources and Tammy Potter, Kentucky State Apiarist with Kentucky Department of Agriculture 

Kentucky Department of Agriculture stakeholders with the Kentucky Pollinator Protection and Monarch Conservation group met on July 20th at the Louisville Zoo and Botanical Gardens. This group meets annually to discuss pollinator conservation topics and projects dedicated to improving pollinator habitat and public knowledge on the plight of pollinators in our state. Members include representatives from a variety of sectors: agriculture, education, transportation right of ways, state and local government, federal government, nonprofits, private and public nature preserves, garden clubs, beekeepers, private businesses, and interested members of the public.  

This July, 65 attendees came together to learn about the Louisville Zoo’s pollinator work and hear updates from various stakeholders including the Office of Kentucky Nature Preserves, the Natural Resources Conservation Service, Kentucky Department of Agriculture, Kentucky Department of Fish and Wildlife Resources, Cave Hill Cemetery, and the Waterfront Botanical Gardens.  

This photo shows participants at the pollinator stakeholders meeting.
Photo: Joe Omielan 

After a tour of the zoo’s gardens, led by Matthew Lahm of the Louisville Zoo, participants in this year’s meeting heard from several experts:  

  • Tony Romano, Office of Kentucky Nature Preserves: Roadside Remnants and Pollinator Habitat 
  • Katie Cody, Office of Kentucky Nature Preserves: Pollinators in the Forest 
  • Michaela Rogers, Kentucky Department of Fish and Wildlife Resources and Shelby Fulton, Office of Kentucky Nature Preserves: State Wildlife Action Plan Insect Update and Partner Engagement 
  • Christy Wampler and Nathan Lind, United States Department of Agriculture Natural Resources Conservation Service: USDA-NRCS Updates 
  • Michael Higgs, Cave Hill Cemetery: Cave Hill is More than a Cemetery 
  • Kat Rivers, Waterfront Botanical Gardens: Planning for Pollinators 

Stakeholders made connections with others pursuing pollinator projects throughout Kentucky and learn about resources offered through various programs highlighted by presenters. Columbia Gas and Roundstone Native Seed Company have worked to convert natural gas rights-of-way to pollinator habitat, and the Transportation Cabinet and the Office of Kentucky Nature Preserves are working on similar projects.  

This year, members of this group also learned about the Pollinator Protection Program Web App, which allows farmers and other chemical applicators to communicate via text or email with beekeepers when a label requires​ communication. This app assists with EPA compliancy and is free to farmers, applicators, landowners, beekeepers, etc. 

This photo shows participants at the pollinator stakeholders meeting.
Photo: Joe Omielan 

This group meets annually and tries to find a new site each year.  Last year, they met at James Audubon State Park.  The Kentucky Pollinator Stakeholders group plans to meet again in Berea Kentucky in 2024, with a date and location to be announced later.  


Michael Rogers, Kentucky Dapartment of Fish and Wildlife Resources.

Michaela Rogers is an Environmental Scientist with the Kentucky Department of Fish and Wildlife Resources. She serves as the Monarch and Pollinator Coordinator for the agency and manages implementation of the Kentucky Monarch Conservation Plan. 

After working for six years with coal companies to establish pollinator habitat on former surface mine sites and working in the queen bee season in the winters, Tammy Horn Potter became Kentucky’s State Apiarist in 2014. She collects honey bee samples for the USDA Honey Bee Health Survey and has worked extensively to improve genetic diversity of queen bees with the Kentucky Queen Bee Breeders Association. She has also worked to improve honey label information with the Kentucky State Beekeepers Association. She is particularly proud of the Kentucky Department of Agriculture Pollinator Protection Plan both for its diversity of stakeholders and for the department’s creation of an app, which coordinates spray information between applicators and beekeepers. 

Landscaping with native plants: how to plan your garden

By Jo Ellen Meyers Sharp

Native plants are gaining in popularity, especially with so many gardeners interested in supporting pollinators and other native insects and wildlife. So how can you landscape with native plants? You first have to plan your yard, and we’ll show you how.

Know your land

Before you can decide which native plants you want in your Kentucky native plant garden, you have to know your growing conditions. Is the area in full sun or shade? Does the soil stay wet or is it dry? Is the soil alkaline or acidic?

This photo shows a soil test submission form.

A good first step is to take a soil test. (You can contact your county extension agent for more information.) This determines the pH of the soil and identifies the minerals and nutrients it has or is missing. The results have recommendations on how to correct any problems. Info like the pH (acid or alkaline soil) also guides your plant selection.

Learn which USDA Zone your plot of land is in. Most of Kentucky is in Zones 6a and 6b. Small sections of the southwest are in USDA Zone 7. These zones tell you which plants are hardy enough to survive our Kentucky winters.

Pay attention to the sun. As the sun moves through the season, the amount of light that hits the ground changes. What starts out sunny in spring could be dense shade by summer as trees leaf out.

Look at the competition other plants may present for your Kentucky native plant garden. Will tree roots be a problem? They win when it comes to sucking up nutrients like water. Plus, digging through them can damage or kill a tree. Some conifers tend to be messy, dropping needles, which can make plants underneath unsightly.

Put pencil to paper or use a landscape app

Make a rough sketch of the shape of the garden you want. Note where sidewalks and structures are and where trees and shrubs are. Is there a water source nearby?

Look at how the light moves through the space – is part of a section in full sun while a portion stays in shade? That garden bed may need a mix of shade- and sun-loving plants.

Start small

Designing and planting a new garden bed can be a lot of work. The biggest job will be the soil preparation. You can always extend a bed every year or two until it’s the size and look you want. Besides saving energy, starting small also conserves cash.

Select your native plants

Some native plants are more garden-worthy than others. Although they may have all the benefits gardeners want in native plants, such as attractive leaves and colorful fruit, they may be aggressive and spread beyond their space. Others may be prone to disease, adding to the maintenance and worries about the plants.

The Kentucky Native Plant Project has a comprehensive listing and descriptions of plants, from spring ephemerals to trees. The Kentucky Native Plant Society has a list of plant suppliers and service providers.

Select perennials that provide a sequence of flowers through the seasons. Start with spring-blooming perennials such as columbine (Aquilegia canadensis), followed by summer-blooming garden phlox (P. paniculata), followed by asters (Symphyotrichum novae-angliae) in fall.

Many native shrubs, such as Viburnum, will have spring flowers, attractive summer foliage, fall color, and fruit that persists into winter. Buttonbush (Cephalanthus occidentalis) has round balls for flowers, colorful fall fruit, and leaves. Winterberry holly (Ilex verticillata), a deciduous holly, has red fruit that stands out in the winter landscape.

For trees, look at the Kentucky coffee bean (Gymnocladus dioicus), river birch (Betula nigra), honeylocust (Gleditsia triacanthos), red maple (Acer rubrum), and white oak (Quercus alba) as good shade tree selections. Flowering dogwood (Cornus florida), Eastern redbud (Cercis canadensis), and downy serviceberry (Amelanchier arbora) are excellent native ornamental trees.

Design tips for a native plant garden

Once you have plants selected, follow a few basic design tips.

  • Avoid planting perennials as singles. Rather, plant them in groups of three or five for the best show.
  • Put tall plants in the back of the bed, with medium-height plants in the middle and short plants in the front.
  • The plant placement can be symmetrical or asymmetrical. Maybe a stand of garden phlox at one end will be balanced at the other with a stand of ornamental grass, such as little bluestem (Schizachyrium scoparium).
  • Texture and form are also important. Flowers last only for a short time compared to leaves and the form of a plant.
  • Make sure plants will fit in the garden space once they’ve reached their mature size. It’s hard to cram a 6-foot-wide shrub in a 4-foot-deep bed. Planting for the mature size reduces long-term maintenance.

Other considerations

A lot of people think native plants will survive with little or no care, but that’s not true. Plants are living things and need a little TLC to thrive, bloom, flower, fruit, and shade as you want.

Water newly planted specimens at least once a week. New plants need roughly 1 inch of water every week to 10 days. Most native perennials do not need a lot of fertilizer. Too much fertilizer or soil that is too rich will cause perennials to flop. During exceptionally dry periods, water trees, shrubs, and perennials.

Soil prep is probably the hardest part of planning a garden, especially if you’re a beginner. You’ll need to remove any grass and weeds and add compost, chopped leaves, or other natural amendments to the soil.

Digging the soil for a new bed may unearth decades-old weed seeds, which will grow once exposed to light and water. Consider bringing in a planter’s mix from a local landscape supply company.

A lot of gardening is trial and error. Sometimes plants work out and sometimes they don’t. Don’t be afraid to pull out what you don’t like and plant something new. The more experience you have the better able you’ll be to select plants that work in your beautiful landscape design of native plants.


Jo Ellen Meyers Sharp

Jo Ellen Meyers Sharp is an award-winning Indianapolis-based garden writer, editor, and speaker. Known as a “hortiholic,” she frequently says her eyes are too big for her yard. She blogs at hoosiergardener.com.